The study carried out in Pakke Tiger Reserve in Arunachal Pradesh noted that hornbills, one among the large-sized frugivores, are the top seed dispersers.
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- The large-seeded trees mainly depended on hornbills and imperial pigeons for their dispersal.
- They swallow the fruit as a whole causing no damage to the seed. They are known to disperse seeds far away from the parent plant up to 13 km parent plant up to 13 km.